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Sharp Claws first level gives + 6 MD and + 10% Melee Strength, 2nd rank of it removes these bonuses and replaces them with a + 12 Melee Attack.
Amphibious Assault rank 1 gives + 20% range, rank 2 gives nothing, rank 3 gives +12% missile damage.
I've got limited experience with skill trees, made a couple of submods, but I think its got to do with not stacking the bonuses in the levels, like with building recruitment where you have to tell the game that a lvl 5 Barracks can still recruit tier 1 infantry.