achivements are where you place your first townhall.

Good info here. Are achievements also based on town hall tiles rather than province biome? Was wondering why I got desert bonuses from a grassland province.

I like to play jungle biome but every time I cant decide first pick. Even if I set all fruit for my foresters, fruit collectors doesnt seem efficent for me, in the other hand mushroom consumes so much wood. What you guys usually pick?

Grassland biome bonuses are out of date. Level 1 => +50% Ranch Productivity or +20 Gold for each claimed province. Level 2 => +50% Hunting Lodge Productivity or +35% Ranch Productivity.

Currently, the biome bonus you receive is based on the actual terrain that your 14x14 Town Hall is built on. This can be confusing because you can see in the zoomed out map view that a particular region says Desert Biome but it actually has desert and grasslands, for example. So it doesn't matter what the big map says the biome type is for that region, it matters what tiles you physically place the Town Hall on. And per Azi's comment below, the understanding is that if you place your Town Hall on multiple biomes, the majority should win out.

yes the biome bonus is base off the highest amount of tiles of the same type, so you can place you city hall on a space that is 1/4 forest, 1/4 desert and 2/4 grassland/savanna and you will get savanna . with that in mind remember some bonuses you get only effect while placed on that tile...like how jungle medicine only works on medical farms on jungle tiles.
my personal pick is grassland as the hunting and ranching bonus is all tiles, and only the grassland roamer is grassland only, so find a spot with some forest for early food(fruit) some grassland (just enough for city hall and maybe for some ranches) and desert near water for clay and irrigation ditches

I think the bome bonus is not based on the specified tile. Instead, it depends on where you build your town center. So building on desert surrounded by forest or jungle will still give you the desert biome bonus.

Can anyone confirm?

Thanks for the summary!

Thanks for the useful info!