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You have to rush to get the second gristmill immediately (with your first 60 food), since you will never have a chance to claim it again once you start getting attacked.
You have a narrow window to get 1-3 hives set up before 1st wave, and setting them on the ramp first is essential, as they'll get free hits in while some enemies focus the gristmill.
Then you have to build a few hives and two fences to the south just in case, a farm or two to accelerate funds, and then start amassing hives at the north ramp. And hoping that they won't send a large army south. And if you ever lose more than a few buildings you won't catch up again unless the enemy is almost starved by then.
The Deadbones rush was super easy thanks to Puff Ball's seed and tactic. Apparently achievements only unlock at Army Size 6 !!
you'll wanna place the squirrels in the mill zone across from the enemy spawn, and you can take pot-shots for free against any spawning spots they set up (so long as they don't spawn anything before they die). after you kill off most of their early stuff with at least 9 squirrels, set down moles so they can take out the enemy main mill and any spawners/defenses they have set up. was fairly nice for me, cause they had lizards and doves as their cheap guys (and I drained them of meat by destroying whatever they put down).